Since 1982 he has been engaged in the fields of taxonomy, ecology. cell development and ontogeny of protozoa; and worked on many ciliate groups from almost all biotopes: free living, commensal or parasitic forms in marine, Antarctica, freshwater and soil habitats. He (usually as senior author) and his colleagues discovered and established many new categories including 5 new families, 13 new genera, more than 50 new species, about 20 new combinations or new names. At the same time, he erected and described morphological modes of 17 genera / species. Representative of his over 50 academic papers published are 1) Taxonomic researches on ciliata of periphyton (Protozoa, Ciliophora) in the Poppelsdorfer Weiher, Bonn (1989); 2) Infraciliature and silverline system of the ciliate Pseudovorticella trepida, n. g., n. comb. and establishment of a new family (1990); Morphology and morphogenesis of the freshwater scuticociliate Uronema nigricans (1991); Morphogenesis of the soil ciliate Bakuella edaphoni n. sp. and revision of the genus Bakuella (1992); Contribution to the morphogenesis of the marine ciliate Diophrys scutum (1993); On two soil ciliates of the genus Dileptus (1994); Taxonomy and ecology of the ciliate fauna in the endopagial and pelagial of the Weddel Sea, Antarctica (1995).

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